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Spedding and his colleagues think that the unique aerodynamic wake signatures of bat wings are caused by different mechanical operations in the upstroke of the wingbeat.

斯佩丁同事们认为，独特气签字后，蝙蝠翅膀所同机械操作，在upstrokewingbeat 。

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原声例句
TED-Ed（视版）

Hummingbirds however, turn their wrists during the upstroke, changing their wing's angle to flap in a figure 8 pattern that continually generates lift throughout their wingbeats.

而，蜂鸟在上击时转动手腕，改变翅膀的角度，以 8 字形模式拍打，在整个拍翅程中不断产生升力。

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PBS地球-动物趣闻

Their wing joints are also stiffer than those of flying birds, which helps them produce thrust on both the downstroke - like most birds do - and on the upstroke of the wing.

它们的翅膀关节也比飞鸟的关节更硬，有助于它们像大多数鸟类一样在下击和上击时产生推力。

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