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In both types of feather, the barbules that extend from the barb ramus grow from a single layer of cells, called the barbule plate, on the periphery of the barb ridge.

而论是何种羽毛，从羽枝主枝延出来的羽小枝，都从羽枝脊表「羽小枝板」的单层细胞长出来。

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原声例句
大英百科全书

Each barb branches into barbules, which link to one another by hooks.

每根支分拨成，通过钩子相互连接。

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Space – Verge Science

And lots of times, there are little feather barbules in there that we can look at the microscopic characters

很多时候， 里面有，我们可以看到微观特征。

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PBS地球-动物趣闻

The central part of the feather became a hardened structure called a rachis, with lots of little barbs and barbules, creating the vanes of the feather.

央部分变成了一种称为轴硬化结构，上面有许多倒钩和枝，形成了叶片。

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