It's the opening of a tube, called the spiracular tract, that's used to bring water in toward the gills.

这是根管子的， 为孔道，用于将水带入鳃中。

In Eusthenopteron, the spiracular tract opens up near the front of the skull, just like in most water-breathing fish with gills.

在 Eusthenopteron 中， 孔在头骨前部附近打，就像大多数带鳃的水肺。

And by the time the Devonian Period had ended and the Carboniferous was underway, they had also lost their spiracular tracts, and started using a totally different kind of skull-hole for breathing, called nostrils.

到泥盆纪结束和石炭纪始时，他们也失去了孔，并始使用种完全不同的颅骨孔来呼吸，为鼻孔。