Particularly when they are engaged in other activities, like copulation, which requires coming partway out of the shell.

尤其是当它们忙于其他活动时，比如交配，这需要它们的部分身体从壳里出来。”

The rituals displayed can convey reproductive fitness or they can just trick reluctant mates into copulation.

所展示的仪式以传达生殖能力，也以只是欺骗不情愿的配偶进行交配。

For males, it might just be a few moments of copulation and that's it.

对于雄性来说，这能只是短暂的交配，仅此而已。

(Laughter) Now, we define extra-pair copulation as the mating outside of a pair bond.

（笑声） 现在，我们将配对之外的交配定义为配对之外的交配。

It originates from the copulation of the dragon and the tiger.

它源自龙虎交合。

Researchers tested to see whether showing several demonstrations of copulation would lead to a higher percentage of learned mate preference.

研究人员进行了测试，以了解展示多次交配示范是否会导致更高比例的习得择偶偏好。

But is there anything above and beyond that that is the result of actual copulation and interacting with those spines?

但除此之外还有什么是实际交配和与这些刺相互作用的结果吗？

For example, the fact that I am really outgoing and I'm not shy talking about sex and copulation and things like that.

例如，我真的很外向，而且我并不害羞地谈论性和交配之类的事情。

And it's obviously not exactly what happens during copulation but it's the closest we're going to get when we look at these large vertebrates.

这显然不是交配过程中发生的事情，但这是我们观察这些型脊椎动物时最接近的情况。

This act of copulation, fertilization perhaps, which should have been no more to me than a bee is to a flower, had become for me indecorous, an embarrassing breach of propriety, which it hadn't been before.

这种交配、受精的行为，也许对我而言，就像蜜蜂对一朵花一样，对我来说已经变得不雅，一种令人尴尬的违反礼仪的行为，这在以前是没有的。