One inclusion formed somewhat later exterior to the exosporium. Another inclusion was formed within the exosporium and remained with the spore after mother cell lysis.

当芽形细裂解后，一种放出来，形自由的伴，另一种在芽外壁内侧形，熟后包裹在芽外壁以内不脱落，始终和芽粘连在一起。