Evolution is regarded as a branching process, whereby populations are altered over time and may speciate into separate branches, hybridize together, or terminate by extinction.
Second, some organisms break the concept's one big rule — they produce offspring with other species, or hybridize, and those hybrid species can reproduce, too.
But in this case, the queens somehow managed to also develop the ability to produce male offspring with the full mesostructured genome instead of a hybridized one.