Homosporous ferns produce sporophytes by intra gametophytic selfing, inter gametophytic mating, or apogamy.Gametangium ontogeny, genetic load, and ploidy affect the mating system of each species.
The ontogeny of epiphyllous microsporangia Ginkgo indicated that the microsporophyll in Ginkgo is of phyllome origin.Possibly, the epiphyllous microsporangia is a peculiar chimera.