There must sometimes be a physical impossibility in the male element reaching the ovule, as would be the case with a plant having a pistil too long for the pollen-tubes to reach the ovarium.

有时雄性成分到达珠在物理上是的， 例如雌蕊太长以至于花粉管无法到达子房的植物就是这情况。

In certain whole groups of plants the ovules stand erect, and in others they are suspended; and within the same ovarium of some few plants, one ovule holds the former and a second ovule the latter position.

在某些整植物， 珠直立，而在另一些植物， 珠则悬空；在少数植物的同一个子房内， 一个珠占据前者的位置， 第二个珠占据后者的位置。

If so, the position of the ovules, even when one is erect and the other suspended within the same ovarium, would follow the selection of any slight deviations in position which favoured their fertilisation, and the production of seed.

如果是这样， 珠的位置， 即使一个直立而另一个悬置在同一个子房内，也将遵循有利于它们受精和子生产的任何轻微位置偏差的选择。