While the placoderms were conquering the seas, things were getting even more crowded on land.

当盾皮鱼征服海洋时， 陆地上东西变拥挤。

For example, the skull of a placoderm had an internal braincase made of endoskeletal bone.

例如，盾皮动物头骨有一个由内骨骼骨制成内部脑壳。

And heterostracans, osteostracans and placoderms have been found in both salty AND fresh water environments.

在咸水和淡水环境中都发现了 heterostracans、osteostracans 和 placoderms。

And scientists have seen signs of those changes in the armor of heterostracans, osteostracans and placoderms.

科学家们已经看到了动物、骨动物和盾皮动物盔甲发生这些变化迹象。

In some placoderms, the covering got up to 5 centimeters thick.

在一些盾皮动物中，覆物厚达 5 厘米。

But it's not clear how armored fish, especially placoderms, were related to either of these groups.

但尚不清楚装甲鱼，尤其是盾皮鱼，与这两个群体有何关系。

Now, one of the most widespread placoderms was Bothriolepis, which was able to live in oceanic environments, estuaries, and freshwater settings.

现在，分布最广盾皮动物之一是 Bothriolepis，它能够生活在海洋环境、河口和淡水环境中。

So armored fish came in a variety of shapes and sizes during the Silurian and Devonian, from tiny Heterostracans to massive Placoderms.

因此，在志留纪和泥盆纪期间， 装甲鱼形状和大小各不相同，从微小鱼到巨大盾皮鱼。

While the eurypterines were busy competing with fish, stylonurines moved inland, to brackish and freshwater ecosystems where there were fewer placoderms and less competition.

当广翅鳍鱼忙于与鱼竞争时，苯丁鱼鱼则向内陆移动， 进入咸水和淡水生态系统， 那里盾皮动物较少， 竞争也较少。

So, we probably have placoderms to thank for the nightmare fuel that followed, a family of giant eurypterines that persisted for nearly 40 million years.

因此， 我们可能要感谢盾皮动物为随后噩梦提供了燃料，这是一个持续了近 4000 万年巨型广翅目动物家族。

But as the Silurian gave way to the Devonian, a dangerous new competitive pressure entered the seas — the armored, jawed fish known as the placoderms.

但随着志留纪让位给泥盆纪，一种危险新竞争压力进入了海洋——被称为盾皮鱼有甲有颚鱼。

But placoderms were also unique among armored fish in having a complete, mineralized internal skeleton – meaning they had both an endoskeleton, like you do, as well as an exoskeleton.

但盾皮鱼在有甲鱼中也是独一无二，因为它们拥有完整、矿化内部骨骼——这意味着它们既有内骨骼，就像你一样， 也有外骨骼。

Others think that most placoderms specifically share common ancestors with ray-finned and lobe-finned fishes, but there are some placoderms that are more closely related to lobe-finned fishes than to other armored fish.

其他人认为大多数盾皮鱼与辐鳍鱼和叶鳍鱼有共同祖先，但也有一些盾皮鱼与叶鳍鱼关系比与其他装甲鱼关系密切。

In the Late Devonian, giant armored jawed fish -- the placoderms -- would come to dominate these seas and force other ocean-dwellers to adapt, flee, or perish.

在泥盆纪晚期，巨大有甲颚鱼——盾皮鱼——将主宰这些海洋并迫使其他海洋生物适应、逃离或灭亡。

But despite their best efforts, these invertebrates couldn't keep up with placoderms, and eurypterines lost over 50 percent of their diversity within the first 10 million years of the Devonian.

但是，尽管他们尽了最大努力， 这些无脊椎动物还是赶不上盾皮动物，而广翅目动物在泥盆纪前 1000 万年内失去了超过 50% 多样性。

But the eurypterines that did survive managed to stage a sort of evolutionary rally, getting bigger, faster, and stronger in the Devonian to cope with seas filled with placoderms that were doing the same.

但是，幸存下来广翅目动物成功地进行了一场进化集会，在泥盆纪变大、快、强壮， 以应对充满同样做同样事情盾皮动物海洋。