The changes were, in all probability, made before 1600.
这些变革

发生在1600年之前。
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法则 
择真题(英语二)
读 Or if they have specific kinds of brains, and whether they were capable of performing certain kinds of trade-off behaviours or pain-related behaviours, learning behaviours that we thought would correlate with the probability of sentience.
或者它们是否有特定种类
大脑,以及它们是否有能力进行某种取舍行为或与疼痛相关
行为,以及我们认为可能与感知相关
学习行为。
美味英语