The ovule is bitegminous and tenuinucellate.The archesporial cell locates under the nucellus epidermis, which develops into fuctional megasporocyte.

孢原细位于珠心表皮，接起大孢子母细的功能。

Tapetum (pl. tapeta) A food-rich layer surrounding the spore mother cells in the anthers of vascular plants.

维管植物内包围孢子母细的富含营养物质的一层结构。