The ovule is bitegminous and tenuinucellate.The archesporial cell locates under the nucellus epidermis, which develops into fuctional megasporocyte.

孢细位于珠心表皮之下，直接起大孢子母细功能。

Tapetum (pl. tapeta) A food-rich layer surrounding the spore mother cells in the anthers of vascular plants.

维管植物花粉囊内包围孢子母细营养物质一层结构。